Date of Award

Fall 2025

Project Type

Thesis

Program or Major

Genetics

Degree Name

Master of Science

First Advisor

Konstantinos Sousounis

Second Advisor

Vicki Jeffers

Third Advisor

Sherine Elsawa

Abstract

Unlike humans, highly regenerative organisms such as salamanders encounter several genotoxic threats from repeated cycles of proliferation, resulting in DNA damage. Understanding the mechanisms by which these organisms resolve DNA damage to maintain genomic integrity during regeneration remains poorly understood. This research evaluated the roles of Eya2, Chk1, and Chk2 in the DNA damage response (DDR) during lens regeneration in the newt, Pleurodeles waltl. Pharmacological inhibitors were used to impair Eya2 and Chk1/2 function throughout regeneration. DNA damage, downstream cell cycle regulators, and proliferation markers were evaluated through histological and western blot analyses. Eya2 inhibition significantly reduces eye and regenerated lens size, increases p-Y142-H2AX, dysregulates CCNB1 and p-Chk1, and subsequently reduces S-phase entry. Conversely, Chk1/2 inhibition reduced eye size, but did not disrupt lens regeneration. p-JNK levels were notably reduced, suggesting a possible interplay in the DDR. Overall, Eya2 was found to be a key regulator of DNA damage and cell cycle progression during regeneration across tissue types and multiple salamander species. These results shed light on the significance of the DDR in regenerative organisms as it may inform future therapies for chronic ocular disease.

Available for download on Friday, November 20, 2026

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