Date of Award

Spring 2024

Project Type

Thesis

Program or Major

Genetics

Degree Name

Master of Science

First Advisor

Vicki Jeffers

Second Advisor

Vicki Jeffers

Third Advisor

David Plachetzki

Abstract

Transcription is well-conserved across eukaryotes, lending itself to countless studies in model organisms. Still, some early-diverging eukaryotes, such as Apicomplexans, continue to deviate from canonical gene expression. One such Apicomplexan is the single-celled parasite, Toxoplasma gondii. Toxoplasma maintains many elements of conventional eukaryotic transcription but demonstrates a relatively short 8-hour replication time. This suggests that transcription is tightly regulated in these parasites, consistent with their ability to differentiate between life stages per the demand of the host environment. Transcription initiation has, however, remained elusive in Toxoplasma. The formation of a preinitiation complex (PIC) has not been characterized as the parasites lack a conserved suite of transcription factors. TATA box-binding proteins (TBPs), as initiators of transcription in other eukaryotes, are an obvious place to start to understand recruitment of the PIC to promoters of transcribed genes. Promoters in Toxoplasma, however, are not well-defined. Additionally, the Toxoplasma genome appears to be void of TATA boxes. This research explored DNA and protein interactions of Toxoplasma TBP1 and aimed to identify TATA-like DNA motifs, identify associated transcription factors, and define the set of genes that are subject to its regulation.

Available for download on Wednesday, June 18, 2025

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